Paleontology (British: palaeontology)[note 1] is the study of prehistoric Prehistory is a term used to describe the period before recorded history. Paul Tournal originally coined the term Pré-historique in describing the finds he had made in the caves of southern France.[citation needed] It came into use in France in the 1830s to describe the time before writing, and the word "prehistoric" was introduced into life Life is a characteristic that distinguishes objects that have signaling and self-sustaining processes (biology) from those that do not, either because such functions have ceased (death), or else because they lack such functions and are classified as inanimate, including organisms' evolution and interactions with each other and their environments (their paleoecology Paleoecology uses data from fossils and subfossils to reconstruct the ecosystems of the past. It involves the study of fossil organisms and their associated remains, including their life cycle, living interactions, natural environment, and manner of death and burial to reconstruct the paleoevironment. Paleoecology's aim is therefore to build the). As a "historical science" it tries to explain causes rather than conduct experiments to observe effects. Paleontological observations have been documented as far back as the 5th century BC Anno Domini and Before Christ (abbreviated as BC or B.C.) are designations used to label years in the Julian and Gregorian calendars. The calendar era to which they refer is based on the traditionally reckoned year of the conception or birth of Jesus, with AD denoting years after the start of this epoch, and BC denoting years before the start of. The science became established in the 18th century as a result of Georges Cuvier Georges Léopold Chrétien Frédéric Dagobert Cuvier was a French naturalist and zoologist. Of working class origins, he belonged to a new class of self-made scholars who worked their way to the top of academe. Cuvier was a major figure in natural sciences research in the early 19th century, and was instrumental in establishing the fields of's work on comparative anatomy Comparative anatomy is the study of similarities and differences in the anatomy of organisms. It is closely related to evolutionary biology and phylogeny, and developed rapidly in the 19th century. Fossils found in China since the 1990s have provided new information about the earliest evolution of animals Animals are a major group of mostly multicellular, eukaryotic organisms of the kingdom Animalia or Metazoa. Their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later on in their life. Most animals are motile, meaning they can move spontaneously and independently. All animals are also, early fish, dinosaurs Dinosaurs are a diverse group of reptiles. They were the dominant terrestrial vertebrates for over 160 million years, from the late Triassic period until the end of the Cretaceous period (about 65 million years ago), when the Cretaceous–Tertiary extinction event caused the extinction of most dinosaur species, except for some birds. The fossil and the evolution of birds and mammals Mammals are a class of vertebrate, air-breathing animals whose females are characterized by the possession of mammary glands while both males and females are characterized by hair and/or fur, three middle ear bones used in hearing, and a neocortex region in the brain. Some mammals have sweat glands, but most do not. Paleontology lies on the border between biology Biology is a natural science concerned with the study of life and living organisms, including their structure, function, growth, origin, evolution, distribution, and taxonomy and geology Geology is the science and study of the physical matter and energy that constitute the Earth. The field of geology encompasses the study of the composition, structure, properties, and history of the planet's physical material, the processes by which it is formed, moved, and changed, the history of life on Earth, and human interactions with the, and shares with archaeology Archaeology, or archeology , is the study of past human societies, primarily through the recovery and analysis of the material culture and environmental data which they have left behind, which includes artifacts, architecture, biofacts and cultural landscapes. Due to the fact that archaeology employs a wide range of different procedures, it can be a border that is difficult to define. It now uses techniques drawn from a wide range of sciences, including biochemistry Biochemistry is the study of the chemical processes in living organisms. It deals with the structures and functions of cellular components such as proteins, carbohydrates, lipids, nucleic acids and other biomolecules. Over the last 40 years biochemistry has become so successful at explaining living processes that now almost all areas of the life, mathematics Mathematics is the study of quantity, structure, space, and change. Mathematicians seek out patterns, formulate new conjectures, and establish truth by rigorous deduction from appropriately chosen axioms and definitions and engineering Engineering is the discipline, art and profession of acquiring and applying technical, scientific, and mathematical knowledge to design and implement materials, structures, machines, devices, systems, and processes that safely realize a desired objective or invention. As knowledge has increased, paleontology has developed specialized subdivisions, some of which focus on different types of fossil Fossils are the preserved remains or traces of animals, plants, and other organisms from the remote past. The totality of fossils, both discovered and undiscovered, and their placement in fossiliferous (fossil-containing) rock formations and sedimentary layers (strata) is known as the fossil record. The study of fossils across geological time, how organisms while others study ecological Paleoecology uses data from fossils and subfossils to reconstruct the ecosystems of the past. It involves the study of fossil organisms and their associated remains, including their life cycle, living interactions, natural environment, and manner of death and burial to reconstruct the paleoevironment. Paleoecology's aim is therefore to build the and environmental history, such as ancient climates Paleoclimatology is the study of climate change taken on the scale of the entire history of Earth. It uses records from ice sheets, tree rings, sediment, corals, shells and rocks to determine the past state of the climate system on Earth.

Body fossils and trace fossils Trace fossils, also called ichnofossils , are geological records of biological activity. Trace fossils may be impressions made on the substrate by an organism: for example, burrows, borings (bioerosion), footprints and feeding marks, and root cavities. The term in its broadest sense also includes the remains of other organic material produced by are the principal types of evidence about ancient life, and geochemical The field of geochemistry involves study of the chemical composition of the Earth and other planets, chemical processes and reactions that govern the composition of rocks and soils, and the cycles of matter and energy that transport the Earth's chemical components in time and space, and their interaction with the hydrosphere and the atmosphere evidence has helped to decipher the evolution of life before there were organisms large enough to leave fossils. Estimating the dates of these remains is essential but difficult: sometimes adjacent rock layers allow radiometric dating Radiometric dating is a technique used to date materials, usually based on a comparison between the observed abundance of a naturally occurring radioactive isotope and its decay products, using known decay rates. It is the principal source of information about the absolute age of rocks and other geological features, including the age of the Earth, which provides absolute dates that are accurate to within 0.5%, but more often paleontologists have to rely on relative dating by solving the "jigsaw puzzles A jigsaw puzzle is a tiling puzzle that requires the assembly of numerous small, often oddly shaped, interlocking and tessellating pieces. Each piece usually has a small part of a picture on it; when complete, a jigsaw puzzle produces a complete picture. In some cases more advanced types have appeared on the market, such as spherical jigsaws and" of biostratigraphy Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them. Usually the aim is correlation, demonstrating that a particular horizon in one geological section represents the same period of time as another horizon at some other section. Classifying ancient organisms is also difficult, as many do not fit well into the Linnean taxonomy The same applies to "Linnaean name": depending on context this may either be a formal name given by Linnaeus , such as Giraffa camelopardalis Linnaeus, 1758, or a formal name in the accepted nomenclature (as opposed to a modernistic clade name) that is commonly used for classifying living organisms, and paleontologists more often use cladistics Cladistics is a method of classifying species of organisms into groups called clades, which consist of 1) all the descendants of an ancestral organism and 2) the ancestor itself. For example, birds, dinosaurs, crocodiles, and all descendants of their most recent common ancestor form a clade. In the terms of biological systematics, a clade is a to draw up evolutionary "family trees". The final quarter of the 20th century saw the development of molecular phylogenetics Molecular phylogenetics, also known as molecular systematics, is the use of the structure of molecules to gain information on an organism's evolutionary relationships. The result of a molecular phylogenetic analysis is expressed in a phylogenetic tree, which investigates how closely organisms are related by measuring how similar the DNA Deoxyribonucleic acid ( /diːˌɒksɨˌraɪbɵ.nuːˈkleɪ.ɪk ˈæsɪd/ (help·info)) (DNA) is a nucleic acid that contains the genetic instructions used in the development and functioning of all known living organisms and some viruses. The main role of DNA molecules is the long-term storage of information. DNA is often compared to a set of is in their genomes In modern molecular biology and genetics, the genome is the entirety of an organism's hereditary information. It is encoded either in DNA or, for many types of virus, in RNA. The genome includes both the genes and the non-coding sequences of the DNA. Molecular phylogenetics has also been used to estimate the dates when species diverged, but there is controversy about the reliability of the molecular clock The molecular clock (based on the molecular clock hypothesis ) is a technique in molecular evolution that uses fossil constraints and rates of molecular change to deduce the time in geologic history when two species or other taxa diverged. It is used to estimate the time of occurrence of events called speciation or radiation. The molecular data on which such estimates depend.

Use of all these techniques has enabled paleontologists to discover much of the evolutionary history of life The evolutionary history of life on Earth traces the processes by which living and fossil organisms evolved. It stretches from the origin of life on Earth, thought to be over 3,500 million years ago, to the present day. The similarities between all present day organisms indicate the presence of a common ancestor from which all known species have, almost all the way back to when Earth Earth is the third planet from the Sun, and the densest and fifth-largest of the eight planets in the Solar System. It is also the largest of the Solar System's four terrestrial planets. It is sometimes referred to as the World, the Blue Planet,[note 6] or by its Latin name, Terra.[note 7] became capable of supporting life, about 3,800 million years ago. For about half of that time the only life was single-celled A microorganism or microbe is an organism that is microscopic (too small to be seen by the naked human eye). The study of microorganisms is called microbiology, a subject that began with Anton van Leeuwenhoek's discovery of microorganisms in 1675, using a microscope of his own design micro-organisms A microorganism or microbe is an organism that is microscopic (too small to be seen by the naked human eye). The study of microorganisms is called microbiology, a subject that began with Anton van Leeuwenhoek's discovery of microorganisms in 1675, using a microscope of his own design, mostly in microbial mats that formed ecosystems An ecosystem consists of all the organisms living in a particular area, as well as all the nonliving, physical components of the environment with which the organisms interact, such as air, soil, water, and sunlight. It is all the organisms in a given area, along with the nonliving factors with which they interact; a biological community and its only a few millimeters thick. Earth's atmosphere The atmosphere of Earth is a layer of gases surrounding the planet Earth that is retained by Earth's gravity. The atmosphere protects life on Earth by absorbing ultraviolet solar radiation, warming the surface through heat retention , and reducing temperature extremes between day and night. Dry air contains roughly (by volume) 78% nitrogen, 21% originally contained virtually no oxygen Oxygen (pronounced /ˈɒksɨdʒɨn/, OK-si-jin, from the Greek roots ὀξύς (acid, literally "sharp", from the taste of acids) and -γενής (-genēs) (producer, literally begetter), is the element with atomic number 8 and represented by the symbol O. It is a member of the chalcogen group on the periodic table, and is a highly, and its oxygenation Oxygen (pronounced /ˈɒksɨdʒɨn/, OK-si-jin, from the Greek roots ὀξύς (acid, literally "sharp", from the taste of acids) and -γενής (-genēs) (producer, literally begetter), is the element with atomic number 8 and represented by the symbol O. It is a member of the chalcogen group on the periodic table, and is a highly began about 2,400 million years ago. This may have caused an accelerating increase in the diversity and complexity of life, and early multicellular Multicellular organisms are organisms that consist of more than one cell. Each cell is specialized to do a certain job for that organism. Most life that can be seen with the naked eye is multicellular, as are all members of the kingdoms Plantae and Animalia plants Plants are living organisms belonging to the kingdom Plantae. They include familiar organisms such as trees, herbs, bushes, grasses, vines, ferns, mosses, and green algae. The scientific study of plants, known as botany, has identified about 350,000 extant species of plants, defined as seed plants, bryophytes, ferns and fern allies. As of 2004, and fungi A fungus ) is a member of a large group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, Fungi (pronounced /ˈfʌndʒaɪ/ or /ˈfʌŋɡaɪ/), that is separate from plants, animals and bacteria. One major difference is that fungal have been found in rocks dated from 1,700 to 1,200 million years ago. The earliest multicellular animal fossils are much later, from about 580 million years ago, but animals diversified very rapidly and there is a lively debate about whether most of this happened in a relatively short Cambrian explosion The Cambrian explosion or Cambrian radiation was the relatively rapid appearance, over a period of many million years, of most major groups of complex animals around 530 million years ago, as found in the fossil record. This was accompanied by a major diversification of other organisms, including animals, phytoplankton, and calcimicrobes. Before or started earlier but has been hidden by lack of fossils. All of these organisms lived in water, but plants and invertebrates An invertebrate is an animal without a backbone. The group includes 95% of all animal species — all animals except those in the chordate subphylum Vertebrata started colonizing land from about 490 million years ago and vertebrates Vertebrates are members of the subphylum Vertebrata, chordates with backbones and spinal columns. About 58,000 species of vertebrates have been currently described. Vertebrata is the largest subphylum of chordates, and contains many familiar groups of large land animals. Vertebrates are comprised of animals from the groups jawless fishs, bony fish, followed them about 370 million years ago. The first dinosaurs Dinosaurs are a diverse group of reptiles. They were the dominant terrestrial vertebrates for over 160 million years, from the late Triassic period until the end of the Cretaceous period (about 65 million years ago), when the Cretaceous–Tertiary extinction event caused the extinction of most dinosaur species, except for some birds. The fossil appeared about 230 million years ago and birds Birds are winged, bipedal, endothermic (warm-blooded), egg-laying, vertebrate animals. There are around 10,000 living species, making them the most varied of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from the 5 cm (2 in) Bee Hummingbird to the 2.75 m (9 ft) Ostrich evolved from one dinosaur group about 150 million years ago. During the time of the dinosaurs, mammals Mammals are a class of vertebrate, air-breathing animals whose females are characterized by the possession of mammary glands while both males and females are characterized by hair and/or fur, three middle ear bones used in hearing, and a neocortex region in the brain. Some mammals have sweat glands, but most do not' ancestors survived only as small, mainly nocturnal insectivores, but after the non-avian dinosaurs became extinct in the Cretaceous–Tertiary extinction event 65 million years ago mammals diversified rapidly. Flowering plants The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Together with gymnosperms, they are the only extant groups of seed-producing plants, but they can be distinguished from the gymnosperms by a series of synapomorphies (derived characteristics). These characteristics include flowers, appeared and rapidly diversified between 130 million years ago and 90 million years ago, possibly helped by coevolution In a broad sense, biological coevolution is "the change of a biological object triggered by the change of a related object". Coevolution can occur at multiple levels of biology: it can be as microscopic as correlated mutations between amino acids in a protein, or as macroscopic as covarying traits between different species in an with pollinating Pollination is the process by which pollen is transferred in plants, thereby enabling fertilization and sexual reproduction. Pollen grains, which contain the male gametes to where the female gamete(s) are contained within the carpel; in gymnosperms the pollen is directly applied to the ovule itself. The receptive part of the carpel is called a insects. Social insects Eusociality is a term used for the highest level of social organization in a hierarchical classification appeared around the same time and, although they have relatively few species, now form over 50% of the total mass of all insects. The upright-walking common ancestor of humans and chimpanzees Sahelanthropus tchadensis appeared around 6 to 7 million years ago, and anatomically modern humans appeared under 200,000 years ago. The course of evolution has been changed several times by mass extinctions that wiped out previously dominant groups and allowed other to rise from obscurity to become major components of ecosystems.

Contents

Definition

A paleontologist carefully chips rock from a column of dinosaur vertebrae.

The simplest definition is "the study of ancient life".[1] Paleontology seeks information about several aspects of past organisms: "their identity and origin, their environment and evolution, and what they can tell us about the Earth's organic and inorganic past".[2]

A historical science

Paleontology is one of the historical sciences, along with archaeology, geology, biology, astronomy, cosmology, philology and history itself.[3] This means that it aims to describe phenomena of the past and reconstruct their causes.[4] Hence it has three main elements: description of the phenomena; developing a general theory about the causes of various types of change; and applying those theories to specific facts.[3]

When trying to explain past phenomena, paleontologists and other historical scientists often construct a set of hypotheses about the causes and then look for a "smoking gun", a piece of evidence which indicates that one of the hypotheses is a better explanation than the others. Sometimes the "smoking gun" is discovered by a fortunate accident during other research, for example the discovery by Luis Alvarez and Walter Alvarez of an iridium-rich layer at the CretaceousTertiary boundary made asteroid impact and volcanism the most favored explanations for the Cretaceous–Tertiary extinction event.[4]

The other main type of science is experimental science, which is often said to work by conducting experiments to disprove hypotheses about the workings and causes of natural phenomena – note that this approach cannot prove a hypothesis is correct, since some later experiment may disprove it. However, when confronted with totally unexpected phenomena, such as the first evidence for invisible radiation, experimental scientists often use the same approach as historical scientists: construct a set of hypotheses about the causes and then look for a "smoking gun".[4]

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